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From Latin ad (to) and nidus (nest).[1]

Coined by geneticist Wilhelm Ludwig in 1950[2][3] as "a fifth evolutionary factor".[4]


annidation (plural annidations)

  1. (ecology) The adaptation of the various genotypes to different ecological niches, such as where a mutant form of a species is maintained in a population because it can flourish in a way that the parent organisms cannot.[5][1]
    • Flax grew more rapidly and matured earlier than linseed and this difference in timing of growth led to annidation.[6]


  1. 1.0 1.1 "annidation." (German) Lexikon der Biologie. Spektrum Akademischer Verlag. 14 Jun. 2009.
  2. ^ Ludwig, W. (1950): Zur theorie der konkurrenz. Neue Ergeb. Prob. Zool., Klatt-Festschrift., 516-537.
  3. ^ Bolnick, Daniel I., (2007): Behavioural Genetics: Evolutionary Fingerprint of the 'Invisible Hand'. Current Biology 17 (15): R597. doi:10.1016/j.cub.2007.05.056 (PDF)
  4. ^ Antonovics, J. (1990): Wilhelm Ludwig and his contributions to population genetics. Trends in Ecol. Evol. 5: 87-90.
  5. ^ "annidation." McGraw-Hill Dictionary of Scientific and Technical Terms. McGraw-Hill Companies, Inc., 2003. 14 Jun. 2009.
  6. ^ Khan, M.A., P.D. Putwain & A.D. Bradshaw (1975): Population interrelationships: Frequency-dependent fitness in Linum. Heredity, 34: 145–163. doi:10.1038/hdy.1975.20 (abstract)